Early Earth - Origin of Life

Chapter 26

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Theories on How Life Originated

Chemical evolution - abiotic synthesis of organic monomers - formation of complex polymers - aggregates of abiotically-produced molecules (protobionts) developed ability to maintain an environment different from surroundings - developed ability to reproduce - led to the origin of life

Panspermia - organic matter reached earth via comets and meteors - evolved into living material

Theories on Where Life Originated

Traditional view is that life originated in shallow water environment or moist sediments

More recently, evidence suggests that life may have originated on ocean bottom near areas of volcanic activity (sea vents)

Matter has not been resolved - both ideas supported by credible evidence

Origin of Hereditary Information

Very significant development - made reproduction possible and allowed natural selection to operate - made Darwinian evolution possible

RNA was apparently the first genetic material to appear - this has been challenged by some who suggest a simpler hereditary mechanism (not involving nucleic acids)

Major Events in History of Life

Origin of life - about 4 billion years ago

Appearance of prokaryotes - 2.5 billion years ago

Photosynthetic prokaryotes released oxygen - changed nature of atmosphere

Appearance of eukaryotes - 2 billion years ago

Appearance of multicellular organisms - 1 billion years ago

Colonization of land by plants and fungi - 400 million years ago

Comparison of Cell Types
Prokaryotic

Few organelles - no endomembrane system

Chromosome circular - occurs in "nucleoid" region (no nucleus)

No cytoskeleton

No mitosis, meiosis, or sex in life cycle

Other differences

Eukaryotic

Numerous organelles - extensive endomembrane system

Chromosomes linear - located in membrane-bound nucleus

Cytoskeleton present

Life cycles include mitosis, meiosis, sex

Three Major Lineages of Life

Bacteria - prokaryotes - decendents include modern bacteria

Archaea - prokaryotes - decendents include organisms known as extreme thermophiles, extreme halophiles & methanogens

Eukarya - eukaryotic - decendents include all eukaryotes

Classification Systems

Two Kingdom - Plantae and Animalia

Five Kingdom - proposed by R. H. Whittaker in 1969 - lumps prokaryotes into Kingdom Monera - many of the primitive eukaryotes into Kingdom Protista - separated Fungi from Plantae

Eight-Kingdom - separated Monera into Bacteria and Archaea - further subdivided Protista

Three-Domain - most recent - established Domain concept (Bacteria, Archaea & Eukarya)

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