Plant Growth and Structure

Chapter 35

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Angiosperms

•      Most diverse and widespread group of plants - 275,000 species worldwide

•      Adaptations to terrestrial environment - root system, shoot system, vascular tissues (xylem & phloem)

•      Two classes - Monocots & Dicots

 

            Characteristics of
    Monocots               Dicots

  • One cotyledon
  • Veins usually parallel
  • Arrangement of vascular bundles complex
  • Fibrous root system
  • Floral parts usually in multiples of 3

 

  • Two cotyledons
  • Veins usually netlike
  • Vascular bundles arranged in ring
  • Taproot usually present
  • Floral parts usually in multiples of 4 or 5

 

 

Root System

•      Tap root - vertical, with lateral roots - serves to anchor plant - present in dicots

•      Fibrous root - lateral, matlike - present in monocots

•      Adventitious - above ground - serves as prop for support

•      Root hairs on both types - increases surface area

 

Shoot System

•      Vegetative - consists of stem and attached leaves

•      Alternating system of nodes (points where leaves attached) and internodes (segment between nodes)

•      Growth usually concentrated at apex - presence of terminal bud usually inhibits growth of axillary buds (apical dominance)

 

Leaves

•      Main photosynthetic organ of most plants

•      Consists of blade & petiole (absent in some monocots)

•      Simple - single undivided blade

•      Compound - blade divided into leaflets

•      Doubly compound - leaflets subdivided

•      Many leaves modified - e.g., cactus spines

 

Cell Types in Plants -
1) Parenchyma

•      Primary cell walls thin and flexible (secondary walls lacking in most)

•      Large central vacuole generally present

•      Perform most of metabolic functions of plant (e.g., photosynthesis)

•      Occur in leaves, stems & roots

•      Most developing cells of this type before becoming specialized

 

Cell Types in Plants -
2) Collenchyma

•      Thicker primary walls - lack secondary walls

•      Usually grouped in strands or cylinders in zones of growth

•      Cells living - lignin absent

•      Provide support without restraining growth

 

Cell Types in Plants -
3) Sclerenchyma

•      Secondary walls usually thick - rigid -  strengthened with lignin

•      At maturity, may lack protoplast (dead cells)

•      Function in support - two types

•      Fibers - usually occur in bundles (hemp)

•      Sclereids - scattered among parenchyma cells - gritty texture (nutshells & seed coat)

 

Cell Types of Plants -
4) Xylem

•      Elongated - rigid - strengthened with lignin - cells dead at maturity

•      Function in water conduction

•      Two types - tracheids & vessel elements

•      What we normally call “wood”

 

Cell Types in Plants -
5) Phloem

•      Chains of tubes formed by sieve-tube members - alive at functional maturity, but lack organelles (e.g., nucleus, ribosomes)

•      Function in conduction of nutrients

•      Serviced by companion cells connected by plasmodesmata

Tissue Systems

•      Dermal tissue (epidermis) - generally single layer - tightly packed - covers & protects

•      Vascular tissue - continuous throughout plant - arranged differently in each organ - functions in water and nutrient transport

•      Ground tissue - located between dermal and vascular tissues - responsible for most metabolic functions

 

Plant Growth

•      Generally “indeterminate” - continues throughout life (note: animals determinate)

•      Meristems - perpetually embryonic tissues - make indeterminate growth possible

•      Primary growth - increase in length - apical meristems (tips of roots and buds of shoots)

•      Secondary growth - increase in girth - lateral meristems (cylinders of cells)

 

Primary Growth

•      Apical meristems extend roots & shoots - give rise to primary plant body

•      Primary meristems (protoderm, procambium & ground meristem) give rise to 3 primary tissues respectively (dermal, vascular & ground)

•      Three zones of growth - zones of cell division, elongation & maturation

 

Secondary Growth

•      Lateral meristems - produce secondary vascular tissue & periderm

•      Vascular cambium - develops into meristematic cylinder - produces secondary xylem & phloem

•      Cork cambium - gives rise to periderm (protective covering)

•      Secondary tissues external to vascular cambium

 

Anatomy of a Leaf

•      Consists of 2 epidermal layers (upper & lower), palisdae parenchyma & spongy parenchyma

•      Openings (stomata) allow entry and exit of water and gases

•      Guard cells regulate opening and closing of stomata

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